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There are some 40,000 indigenous peoples of the Fly River drainage in Papua New Guinea. The 4,000-mm rainfall contour ecologically demarcates hunter-horticulturalist peoples living in the rainforests of the Upper Fly from hunter-gatherer peoples living in the savanna-swamplands of the Middle and Lower Fly. A complex of factors operate to create significant physical differences between Upper Fly peoples and those of the Middle and Lower Fly. The ecological division between rainforests and savanna-swamplands demarcates a clear clinal separation by stature of Upper Fly peoples from those of the Middle and Lower Fly.
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